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Lichenicolous Fungi – Worldwide
Checklist Including Isolated Cultures and
Sequences Available Phylogenetic Arrangement of Genera |
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Major fungal culture collections that contain lichenicolous strains Cited literature on lichenicolous
fungi
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Species (lichenicolous
fungi in bold; |
Cultures (with accession
numbers when known; culture
collection links) |
Sequences (with accession
numbers) |
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ASCOMYCOTA |
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ARTHONIOMYCETES |
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Arthoniales |
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Arthoniaceae |
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Arthonia
s. lat. [incl. anam. Helicobolomyces
and Subhysteropycnis |
abrothallina Nyl.: on Pseudocyphellaria agelastica R. C. Harris & Lendemer: on Lecanora louisiana almquistii Vain.: on Amygdalaria, Koerberiella, Porpidia
and Trapelia amylospora Almq.: on Porpidia and Farnoldia anatolica Halıcı & Candan: on Aspicilia contorta anjutii S. Y. Kondr. & Alstrup: on Teloschistes velifer apotheciorum (A. Massal.) Almq.: on Myriolecis albescens arthoniicola Diederich & Aptroot: on Arthonia catenulata aspiciliae (Metzl.) S. Y. Kondr.: on Aspicilia atropunctata
Vain.: on sterile foliicolous cf. Echinoplaca ayseniae Halıcı & Candan: on Acarospora [as ‘aysenae’] badia Wedin & Hafellner: on Pseudocyphellaria
granulata biatoricola Ihlen & Owe-Larss.: on Biatora efflorescens caerulescens (Almw.) R. Sant.: on Lecanora varia calicii Kantvilas & Wedin: on Calicium tricolor [as
‘caliciae’] catillarioides Etayo: on Sticta ceracea Etayo & Breuss: on Peltigera polydactylon
[a homonym of Arthonia ceracea Fée] cinnabarinula Müll. Arg.: on foliicolous Porina and Trichothelium
[anamorph: Helicobolomyces lichenicola Matzer] circinata Th. Fr.: on Umbilicaria clemens (Tul.) Th. Fr.: on Rhizoplaca cohabitans Coppins: on Arthothelium reagens colombiana Etayo: on Cladonia coriifoliae Wedin & Hafellner: on Pseudocyphellaria
coriifolia coronata Etayo: on Flavoparmelia caperata cryptotheciae
Matzer & R. Sant.: on foliicolous Cryptothecia
candida darbishirei Follmann & Werner: on Roccella nigerrima destruens Rabenh.: on Physcia destruens var. nana Grube
& Hafellner: on Xanthoria fallax digitatae Hafellner: on Cladonia digitata digitispora Etayo: on Sticta [as ‘digitospora’] diorygmae S. Joshi, D. K. Upreti & S. Nayaka: on Diorygma diploiciae Calat. & Diederich: on Diploicia canescens epicladonia (Nyl.) Alstrup & Zhurb.: on Cladonia epifarinosa Etayo: on Pannaria epimela (Norman) I. M. Lamb: on Buellia punctata epiphyscia Nyl.: on Physcia epitoninia Halıcı & Candan: on Toninia excentrica Th. Fr.: on Lepraria and Leprocaulon farinacea (H. Olivier) R. Sant.: on Ramalina farinacea flavicantis Wedin & Hafellner: on Pseudocyphellaria
flavicans follmaniana Diederich: on Roccella fuscopurpurea (Tul.) R. Sant.: on Peltigera galactinaria Leight.: on Myriolecis garajonayi Etayo: on Porina gelidae R. Sant.: on Placopsis gelida glacialis Alstrup & E. S. Hansen: on Rhizoplaca
melanophthalma glaucomaria (Nyl.) Nyl.: on Lecanora rupicola graphidicola Coppins: on Graphis scripta griseopruinosa F. Berger & E. Zimm.: on Pertusaria
pseudocorallina haematommatum Kalb & Hafellner: on Haematomma haematostigma Grube & Hafellner: on Ramalina hawksworthii Halıcı: on Dimelaena hertelii (Calat., Barreno & V. J. Rico) Hafellner & V.
John: on Aspicilia heterodermiae Etayo: on Heterodermia hodgesii Lendemer & R. C. Harris: on Graphis lineola Mycopegrapha hymeniicola Vain.: on Lecanora
[a poorly known species] ingaderiae Follmann: on Ingaderia insularis
Kantvilas & Wedin: on Caloplaca eos intermedia Matzer: on foliicolous Phyllobiale and Porina intexta Almq.: on Lecidella invadens Coppins: on Schismatomma japewiae Grube & Holien: on Japewia lecanoricola Alstrup & Olech: on Lecanora populicola [as
‘lecanoriicola’] lecanorina (Almq.) R. Sant.: on Lecanora albella lepidiota (H. Olivier) R. Sant.: on Fuscopannaria praetermissa lepidophila (Anzi) Clauzade, Diederich & Cl. Roux (comb.
inval.): on Cladonia lobariellae Etayo: on Lobariella lobothalliae Etayo: on Lobothallia alphoplaca maculiformis Wedin & Hafellner: on
Pseudocyphellaria glabra and P. homoeophylla marginalis Etayo: on Pseudocyphellaria mazosiicola Matzer & R. Sant.: on foliicolous Mazosia minuta Wedin & Hafellner: on Pseudocyphellaria
coriifolia molendoi (Frauenf.) R. Sant.: on Caloplaca and Xanthoria navasiana Etayo: on Lepraria nephromiaria Nyl.: on Nephroma nideri (J. Steiner) Clauzade, Diederich & Cl. Roux: on
Caloplaca gr. variabilis obscurior Triebel: on Pilophorus dovrensis oligospora Vězda: on Myriolecis crenulata pantherina Etayo: on Pertusaria pustulata parietinaria Hafellner & A. Fleischhacker: on Xanthoria
parietina parmeliae (C. W. Dodge & G. E. Baker) D. Hawksw. &
Iturr.: on Physcia caesia peltigerea Th. Fr.: on Peltigera and Solorina peltigerina (Almq.) H. Olivier: on Peltigera and
Solorina pelvetii (Hepp) H. Olivier: on Pseudocyphellaria aurata pepei Etayo & Pérez-Ortega: on Parmelina phaeophysciae Grube & Matzer: on Phaeophyscia plectocarpoides (S. Y. Kondr. & D. J. Galloway) Wedin & S. Y.
Kondr.: on Pseudocyphellaria scabrosa polia Etayo & R. Sant.: on Diploicia canescens prominens Follmann: on Ingaderia gracillima protoparmeliae Etayo: on Protoparmelia badia pseudopegraphina Matzer: on foliicolous Mazosia punctella Nyl.: on Diplotomma rangiformicola Brackel & Etayo: on Cladonia rangiformis rimeliicola Diederich: on Parmotrema rinodinicola Candan & Halıcı: on Rinodina oleae rubescens (Arnold) Clauzade, Diederich & Cl. Roux: on
Buellia epipolia rufidula (Hue) D. Hawksw., R. Sant. & Řvstedal: on Umbilicaria sagenedii Vězda & Kantvilas: on Sagenidium molle sampaianae (Diederich & Etayo) Ertz & Diederich: on Fuscopannaria
sampaiana santessoniana Wedin & Hafellner: on Pseudocyphellaria
scabrosa santessonii Matzer: on foliicolous Porina semi-immersa Wedin & Hafellner: on
Pseudocyphellaria sherparum Grube & Matzer: on Lecanora sherparum squamarinae Etayo: on Squamarina lentigera stevensoniana R. C. Harris & Lendemer: on Haematomma accolens stictaria Nyl.: on Pseudocyphellaria stipitata M. M. E. Alves, Aptroot & M. Cáceres: on
unidentified sterile, sorediate, corticolous lichen (?Bacidina, Fellhanera) subaggregata Wedin & Hafellner: on Pseudocyphellaria hirsuta subconveniens Nyl.: on Lobaria subfuscicola (Linds.) Triebel: on Lecanora albella and L.
carpinea subvelutinae (Vain.) R. Sant.: on foliicolous Sporopodium sytnikii S. Y. Kondr.: on Xanthoria tavaresii Grube & Hafellner: on Anthracothecium and Pyrenula tetraspora S. Y. Kondr. & Kärnefelt: on Caloplaca
chilensis thelotrematis Coppins: on Thelotrema tremelloides Etayo: on Heterodermia triebeliae Zhurb.: on Dactylina urceolata (Elenkin) Calat., Barreno & V. J. Rico: on
Aspicilia varians (Davies) Nyl.: on Lecanora rupicola and L.
helicopis versoeensis Alstrup: on Buellia punctata [= A. aggregata
Alstrup, M. S. Christ. & Hafellner] xanthoparmeliarum
Etayo: on Xanthoparmelia |
Alstrup 1993a Alstrup & Cole 1998 Alstrup & Hansen 2001 Alstrup & Olech 1996 Aptroot et al. 1995 Aptroot et al. 1997 Berger & Zimmermann 2016 Brackel 2015 Calatayud et al. 1995 Calatayud et al. 2004a Candan & Halıcı 2009 Diederich 1995 Ertz et al. 2005 Etayo 1993 Etayo 1996 Etayo 2002a Etayo 2010b Etayo 2017 Etayo & Breuss 1998 Etayo & Pérez-Ortega 2016 Etayo & Sancho 2008 Fleischhacker et al. 2016 Follmann & Werner 2003 Grube & Matzer 1997 Grube et al. 1995 Halıcı 2008 Halıcı & Candan 2009 Halıcı & Candan 2011 Hawksworth 1991a Hawksworth & Iturriaga 2006 Hertel 1969 Joshi et al. 2013 Kantvilas & Wedin 2015 Kondratyuk 1996a Kondratyuk & Zelenko 2002 Lendemer et al. 2016 Matzer 1996 Santesson 1993a Santesson et al. 2004 Triebel et al. 1991 Wedin & Kondratyuk 1997 |
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apotheciorum (UPS:Frisch 11/Se23): RPB2: KJ851148; mtSSU: KJ850970 biatoricola (UPS:Thor 24350): RPB2: KJ851149; mtSSU: KJ850990 graphidicola (UPS:Frisch 10/Jp102): RPB2:
KJ851034; mtSSU: KJ850980 molendoi (UPS:Frisch 11/Se36): nrLSU: KJ851051; RPB2: KJ851117; mtSSU: KJ851000 neglectula (UPS:Frisch 10/Se91): nrLSU: KJ851037; RPB2: KJ851118; mtSSU: KJ850989 parietinaria: nrLSU sequences reported in Fleischhacker et al. (2016) for four
specimens peltigerina (UPS:Westberg Frisch 11/Se46): RPB2:
KJ851122; mtSSU: KJ850998 phaeophysciae (UPS:Ertz 11/2): nrLSU: KJ851067; RPB2: KJ851112 stereocaulina (UPS:Westberg Frisch 11/Se48): nrLSU: KJ851062; mtSSU: KJ850999 subfuscicola (UPS:Frisch 11/Se15): RPB2: KJ851111; mtSSU: KJ850972 subfuscicola (UPS:Thor 11/1): RPB2: KJ851110; mtSSU: KJ850971 |
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Briancoppinsia |
cytospora (Vouaux) Diederich, Ertz, Lawrey & van den Boom:
on parmelioid lichens, Lecanora
conizaeoides, Cladonia and Lepra |
Diederich et al.
2011b |
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Lecanographaceae |
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Lecanographa |
imitans Werner & Follmann: on Roccella humboldtiana |
Egea & Torrente
1994 |
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Phacographa |
glaucomaria (Nyl.) Hafellner:
on Lecanora rupicola s. lat. (incl. L. bicincta) |
Hafellner 2009 |
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protoparmeliae (UPS Westberg and Frisch 11Se/49): nrLSU: KJ851075; RPB2:
KJ851137 zwackhii (UPS Frisch 11/Se3):
nrLSU: KJ851048; mtSSU: KJ851021 |
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Plectocarpon |
aequatoriale Etayo: on Sticta arthonioides Diederich: on Lobaria
isidiophora |
Aptroot et al. 1997 Ertz & van den
Boom 2012 Etayo 2017 Joshi et al. 2016b Kukwa et al. 2012 |
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nephromeum (UPS:
Nordin 5813): RPB2: KJ851139; mtSSU: KJ851004 |
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Opegraphaceae |
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Opegrapha s. lat. |
agelaea Fée: on pyrenocarpous lichen cladoniicola Ertz & Diederich: on Cladonia maligna Triebel: on Lecidea fuscoatra stereocaulicola Alstrup & D. Hawksw.: on Stereocaulon |
Alstrup & Hawksworth
1990 Clauzade et al. 1989 Etayo 2017 Kondratyuk et al.
2015 Kondratyuk et al.
2016b Zhurbenko 2013a Zhurbenko &
Vershinina 2014 |
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Paralecanographa |
grumulosa
(Dufour) Egea & Torrente [incl. L.
hemisphaerica (Laundon) Egea & |
Egea & Torrente
1994 |
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grumulosa (Ertz 13565): nrLSU: HQ454555; RPB2: HQ454695 grumulosa (Ertz 13558): nrLSU: HQ454554; RPB2: HQ454694 grumulosa (Tehler 9031): nrLSU: HQ454553; RPB2: HQ454693 grumulosa (Tehler 9010): nrLSU: HQ454552; RPB2: HQ454692 grumulosa (Tehler 9809): nrLSU: HQ454542; RPB2: HQ454682 |
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Roccellaceae |
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Enterographa |
bagliettoae F. Seavey & J.
Seavey: on Bagliettoa epigraphis Etayo &
Sipman: on Graphis epiphylla (Sérus.) Ertz, Diederich
& Sparrius: on foliicolous Coenogonium
flavicans mazosiae Matzer & R. Sant.:
on foliicolous Mazosia osagensis C. A. Morse: on Bacidina punctata
Ertz & Diederich: on Lobaria |
Ertz el. 2005 Etayo 2017 Matzer 1996 Morse 2013 Seavey et al. 2017 |
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Mazosia |
adelphoparasitica Matzer:
on foliicolous Mazosia |
Matzer 1996 |
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Sigridea |
labyrinthica (Follmann) Ertz & Diederich: on Roccella portentosa |
Ertz et al. 2005 |
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Incertae sedis |
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Arthophacopsis |
parmeliarum Hafellner: on Parmelia s. str. |
Hafellner 1998a |
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Arthothelium |
hymeniicola Ertz & Friday:
on Bacidia (hymenium) |
Ertz & Fryday
2017 |
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Cresponea |
lichenicola Aptroot & M. Cáceres: on Pyrenula |
Cáceres et al. 2014 |
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Mixtoconidium |
canariense Etayo: on Ramalina insidiens (Vouaux) Etayo & van den Boom: on Ramalina
fraxinea nashii Hafellner: on Niebla robusta |
Ertz & Diederich
2007 Etayo 1995a van den Boom &
Etayo 2017 |
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Paradoxomyces |
nymanii Matzer: on
foliicolous Strigula |
Matzer 1996 |
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Perigrapha |
lobariae Zhurb.: on Lobaria
japonica agg. |
Ertz et al. 2005 Pérez-Vargas et al.
2013 Zhurbenko et al.
2015a |
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Phacothecium |
varium (Tul.) Trevis.: on
Xanthoria |
Hafellner 2009 |
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Schismatomma s. lat. |
physconiicola Ertz & Diederich: on Physconia |
Ertz & Diederich
2006 |
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Trichophyma |
bunchosiae Rehm: on
foliicolous Strigula similis Matzer: on
foliicolous Strigula |
Matzer 1996 |
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Lichenostigmatales |
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Phaeococcomycetaceae |
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Etayoa |
trypethelii (Flakus & Kukwa) Diederich & Ertz: on
Graphidaceae, Lecanora, Pertusaria, Trypethelium and
other crustose lichens |
Ertz et al. 2014b |
trypethelii: Cultures obtained by Ertz et al. (2014) |
trypethelii (Mukherjee): nrLSU: KF176943; mtSSU: KF176970 trypethelii (Common 9434K): nrLSU: KF176942; mtSSU: KF176969 trypethelii (Common 9215P): nrLSU: KF176941; mtSSU: KF176968 trypethelii (Common 9200G): nrLSU: KF176940; mtSSU: KF176967 |
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Lichenostigma |
alpinum (R. Sant., Alstrup & D. Hawksw.) Ertz &
Diederich: on Lepra, Ochrolechia,
Varicellaria and other lichens [incl. Diederimyces fuscideae
Etayo, D. microsporus Etayo, Phaeosporobolus minutus Etayo] chlaroterae (F. Berger & Brackel) Ertz & Diederich: on Lecanora and other lichens fellhanerae (R. C. Harris & Lendemer) Ertz & Diederich: on
Fellhanera hyalosporum Kalb & Hafellner: on Haematomma eremaeum maureri Hafellner: on Usnea and other macrolichens |
Ertz et al. 2014b Etayo 1995b Etayo & Sancho
2008 Hawksworth &
Hafellner 1986 Kalb et al. 1995 |
chlaroterae: Cultures obtained by Ertz et al. (2014) |
alpinum (Ertz 17519): nrLSU: KF176946;
mtSSU: KF176973 alpinum (Ertz 17522): nrLSU: KF176945;
mtSSU: KF176972 alpinum (Diederich 17379): nrLSU: KF176944; mtSSU: KF176971 chlaroterae (Goward 12-42): nrLSU: KF176949;
mtSSU: KF176977 chlaroterae (Neuberg): nrLSU: KF176948;
mtSSU: KF176975 chlaroterae (Diederich 17329): nrLSU: KF176947;
mtSSU: KF176974 chlaroterae (Goward 12-30): mtSSU: KF176976 maureri (Diederich 17337-teleomorph): nrLSU: KF176954; mtSSU: KF176980 maureri (Diederich 17337-anamorph): nrLSU: KF176952 maureri (Diederich 17326): nrLSU: KF176953;
mtSSU: KF176979 maureri (Diederich 17306): nrLSU: KF176951;
mtSSU: KF176978 |
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Species (lichenicolous
fungi in bold; |
Cultures (with accession
numbers when known; culture collection links) |
Sequences (with accession
numbers) |
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ASCOMYCOTA |
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DOTHIDEOMYCETES |
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Abrothallales |
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Abrothallaceae |
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Abrothallus |
acetabuli
Diederich: on Pleurosticta acetabulum bertianus
De Not.: on Melanelixia boomii
Pérez-Ortega & Suija: on Nephroma brattii
(S. Y. Kondratyuk) Suija &
Pérez-Ortega: on Pseudocyphellaria faveolata bryoriarum
Hafellner: on Bryoria buellianus
De Not.: on Parmelina caerulescens
Kotte: on Xanthoparmelia canariensis
Pérez-Ortega, van den Boom & Suija: on Pseudocyphellaria
aurata cetrariae
Kotte: on Platismatia glauca chrysanthus
Stein: on Usnea cladoniae
R. Sant. & D. Hawksw.: on Cladonia doliiformis
Pérez-Ortega & Suija: on Sticta eriodermae
Suija, Etayo & Pérez-Ortega: on Erioderma ertzii
Suija & Pérez-Ortega: on Lobaria
pulmonaria etayoi
Pérez-Ortega & Suija: on Sticta granulatae
Wedin: on Pseudocyphellaria halei Pérez-Ortega, Suija, D. Hawksw. & R. Sant.: on
Lobaria heterodermiicola Etayo & F.
Berger: on Heterodermia hypotrachynae
Etayo & Diederich: on Hypotrachyna kamchatica (Zhurb. &
Stepanchikova) Pérez-Ortega & Suija: on Nephroma macrosporus Etayo & R. Sant.: on Durietzia microspermus
Tul.: on Flavoparmelia caperata nephromatis
Suija & Pérez-Ortega: on Nephroma niger Etayo: on Everniastrum,
and possibly Parmotrema and Usnea parmeliarum
(Sommerf.) Nyl.: on Parmelia parmotrematis
Diederich: on Parmotrema peyritschii
(Stein) Kotte: on Vulpicida pezizicola
Diederich & R.C. Harris: on Cladonia
peziziformis prodiens
(Harm.) Diederich & Hafellner: on Hypogymnia puntilloi
Brackel: on Lobaria pulmonaria ramalinae
Diederich: on Ramalina secedens
Wedin & R. Sant.: on Pseudocyphellaria stereocaulorum
Etayo & Diederich: on Stereocaulon stictarum
Etayo: on Sticta stroblii
Hafellner: on Menegazzia terebrata suecicus
(Kirschst.) Nordin: on Ramalina teloschistis
Brackel, Pérez-Ortega & Suija: on Seirophora
and Teloschistes tulasnei
M.S. Cole & D. Hawksw.: on Xanthoparmelia usneae
Rabenh.: on Usnea usneae
var. tetraspora Etayo & Osorio: on Usnea welwitschii |
Brackel 2015 Brackel &
Puntillo 2016 Diederich 1990 Diederich et al.
2017 Etayo 2017 Hafellner et al.
2008 Hawksworth 1990a Ihlen & Wedin
2008 Pérez-Ortega et al.
2014 Suija et al. 2015a Zhurbenko et al.
2012 |
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acetabuli (SPO308): ITS: KF816165; nrLSU: KF816232; nrSSU:
KF816215; mtSSU: KF816205; RPB2:
KF816194; RPB1:
KF816192; TEF1:
KF816188 acetabuli (von Brackel 5661): ITS: KP165100 boomii (van den Boom 43228): ITS: KP165092; TEF1: KP165108 buellianus (SPO303): ITS:
KF816166; nrLSU: KF816234; nrSSU:
KF816217; mtSSU:
KF816207; RPB2:
KF816195; RPB1:
KF816191; TEF1:
KF816186 caerulescens (SPOc80):
ITS: HQ702852 canariensis (van den Boom 37849): ITS: KP165087; TEF1: KP165123 cladoniae (AB53): ITS: KF816173; nrLSU: KF816228; nrSSU: KF816220; RPB2: KF816200; RPB1:
KF816190; TEF1:
KF816181 doliiformis (S309):
ITS: KP165086 eriodermae (AB60):
ITS: KP165101; TEF1: KP165126 eriodermae (AB74):
TEF1: KP165117 ertzii (Ertz
12517): ITS: KP165102;
TEF1: KP165127 etayoi (Perez-Ortega
2003): ITS: KP165085;
TEF1: KP165112 granulatae (Perez-Ortega
3166): ITS: KP165094;
TEF1: KP165122 hypotrachynae (SPO302):
ITS: KF816167;
nrLSU: KF816233; nrSSU: KF816218; mtSSU: KF816208; RPB2:
KF816196 kamchatica (S310):
ITS: KP165091; TEF1: KP165107 nephromatis (Spribille
10581): ITS: KP165088;
TEF1: KP165130 nephromatis (AB77):
ITS: KP165105; TEF1: KP165119 nephromatis (Ertz
12514): ITS: KP165103;
TEF1: KP165118 nephromatis (Perez-Ortega
3159): ITS: KP165089;
TEF1: KP165115 nephromatis (Diederich
12530): ITS: KP165104 parmeliarum (AB36): ITS: KF816172; nrLSU: KF816229; nrSSU: KF816222; RPB2:
KF816201; TEF1: KF816182 parmeliarum (AB57): ITS: KF816178; nrLSU: KF816227; nrSSU: KF816221; mtSSU:
KF816211; RPB2:
KF816198; TEF1:
KF816180 parmotrematis (AB1): ITS: KF816176; nrLSU: KF816231;
nrSSU: KF816225; mtSSU: KF816214; RPB2:
KF816204; TEF1:
KF816185 secedens (SPO305): ITS: KF816169; nrLSU: KF816236; nrSSU: KF816216; mtSSU: KF816206 secedens (Perez-Ortega
3158): ITS: KP165098;
TEF1: KP165121 secedens (Perez-Ortega
3156): ITS: KP165095;
TEF1: KP165111 secedens (Perez-Ortega
3161): ITS: KP165093;
TEF1: KP165110 secedens (Perez-Ortega
3162): ITS: KP165097 secedens (Perez-Ortega
3175): ITS: KP165096 sp. (Perez-Ortega 3155): ITS: KP165099; TEF1: KP165124 sp. (von Brackel 5491): ITS: KP165106; TEF1: KP165120 sp. 1 (SP-2014): ITS: KF816168; nrLSU: KF816235;
nrSSU: KF816219; mtSSU: KF816209; RPB2:
KF816197; RPB1:
KF816193; TEF1:
KF816187 sp. 2
(SP-2014): ITS: KF816175; nrLSU: KF816230; nrSSU:
KF816224; mtSSU:
KF816213; RPB2:
KF816203; TEF1:
KF816184 suecicus (AB56): ITS: KF816177; nrLSU: KF816226;
mtSSU: KF816210;
RPB2: KF816199; TEF1:
KF816179 suecicus (SPO304):
ITS: KF816171 usneae (AB20): ITS: KF816174; nrSSU: KF816223; mtSSU: KF816212; RPB2: KF816202; RPB1:
KF816189; TEF1: KF816183 usneae (SPO306):
ITS: KF816170 welwitschii (Perez-Ortega
3171): TEF1: KP165131 welwitschii (AB61):
TEF1: KP165129 welwitschii (Ertz
1083): TEF1: KP165128 welwitschii (Perez-Ortega
3169): ITS: KP165090;
TEF1: KP165116 welwitschii (Perez-Ortega
3168): TEF1: KP165114 welwitschii (Perez-Ortega
3176): TEF1: KP165113 welwitschii (Spribille
24949): TEF1: KP165109 |
|
Lichenoconiaceae |
|
|
|
|
|
Lichenoconium |
aeruginosum Diederich, M. Brand, van den
Boom & Lawrey: on Cladonia cargillianum (Linds.) D. Hawksw.: on Letharia, Parmotrema,
Platismatia, Ramalina and Usnea christiansenii M. S. Cole & D. Hawksw.: on Nodobryoria
abbreviata echinosporum D. Hawksw.: on Cladia muelleri edgewoodensis Alstrup & M. S. Cole: on Parmelia sulcata erodens M. S. Christ. & D. Hawksw.: on various lichens follmannii S. Y. Kondr. & D. J. Galloway: on Lobaria
pulmonaria laevisporum Kalb & Hafellner: on Haematomma lecanorae (Jaap) D. Hawksw.: on various lichens lichenicola (P. Karst.) Petr. & Syd.: on Physcia plectocarpoides S. Y. Kondr. & D. J. Galloway: on Pseudocyphellaria pyxidatae (Oudem.) Petr. & Syd.: on Cladonia reichlingii Diederich: on Ramalina fraxinea usneae (Anzi) D. Hawksw.: on various lichens xanthoriae M. S. Christ.: on Xanthoria, also reported from
Caloplaca, Cetraria, Cetrelia, Heterodermia, Melanelia,
Physcia and Teloschistes |
Alstrup & Cole
1998 Christiansen 1956 Christiansen 1980 Cole &
Hawksworth 2004 Diederich 1986 Hawksworth 1977b Hawksworth 1981 Kalb et al. 1995 Kondratyuk &
Galloway 1995b Kondratyuk et al.
1994 Lawrey et al. 2011 |
aeruginosum: CBS 129239 erodens: Culture obtained
by Crittenden et al. (1995) erodens: CBS 128028 erodens: CBS 128030 erodens: CBS 128704 lecanorae: Culture obtained
by Crittenden et al. (1995) lecanorae: CBS 128024 sp.: CBS 129139 usneae: CBS 128020 |
aeruginosum (CBS
129239, JL359-09): nrLSU: HQ174269; mtSSU: HQ174268 erodens (JL363-09): nrLSU: HQ174267; mtSSU: HQ174266 lecanorae (CBS 128024): nrLSU: HQ174263; mtSSU: HQ174262 usneae (CBS 128020): nrLSU: HQ174265; mtSSU: HQ174264 |
|
Asterinales |
|
|
|
|
|
Genera
incertae sedis |
|
|
|
|
|
Buelliella |
colombiana Etayo: on Phlyctis
andensis dirinariae Diederich: on Dirinaria
picta eximia Kalb &
Hafellner: on Pyxine cocoes inops (Triebel &
Rambold) Hafellner: on Caloplaca heppiae van den Boom: on Heppia
arenacea minimula (Tuck.) Hafellner:
on Pertusaria physciicola Poelt &
Hafellner: on Phaeophyscia and Physcia poetschii Hafellner: on Endocarpon protoparmeliopseos Etayo &
Pérez-Ortega: on Protoparmeliopsis pusilla Hafellner: on Brigantiaea
leucoxantha trypethelii (Tuck.) Hafellner:
on Trypethelium |
Aptroot et al. 1997 Ertz & Diederich
2015 Etayo 2002a Hafellner 1979 Hafellner 1985b Hafellner et al.
2008 Kalb 1990 Pérez-Ortega &
Etayo 2010 Suija & Alstrup
2004 |
|
physciicola (Ertz
19173): nrLSU: KP456148;
mtSSU: KP456173 physciicola (Ertz
18113): nrLSU: KP456147;
mtSSU: KP456172 |
|
Hemigrapha |
asteriscus (Müll. Arg.) D. Hawksw.: on Peltigera atlantica Diederich & Wedin: on Sticta canariensis nephromatis Wedin & Diederich: on Nephroma australe phaeospora Matzer & Lücking: on foliicolous Strigula
phyllogena pilocarpacearum M. Cáceres & Lücking: on Byssoloma and Fellhanera pseudocyphellariae Diederich & Wedin: on Pseudocyphellaria argyracea strigulae Matzer: on foliicolous Strigula tenellula (Müll. Arg.) Matzer: on foliicolous
Porina |
Cáceres &
Lücking 2000 Diederich &
Wedin 2000 Eriksson et al. 2001 Ertz & Diederich
2015 Hawksworth 1975b Matzer 1996 |
|
asteriscus (Ertz
18690): mtSSU: KP456176 |
|
Karschia |
alpicolae D. Hawksw. & Alstrup: on Allantoparmelia alpicola santessonii Hafellner: on Porpidia talcophila (Flot.) Koerb.: on Diploschistes |
Alstrup &
Hawksworth 1990 Ertz & Diederich
2015 Hafellner 1979 Santesson 1994a |
|
talcophila (Diederich
16749): nrLSU: KP456155;
mtSSU: KP456181 |
|
Labrocarpon |
canariense (D, Hawksw.) Etayo & Pérez-Ortega: on
Pertusaria |
Ertz & Diederich
2015 Hawksworth 1982b Pérez-Ortega &
Etayo 2010 |
|
canariense (Ertz 16907): nrLSU: KP456158; mtSSU: KP456184 |
|
Melanographa |
tribuloides (Tuck.) Müll. Arg.: on Anthracothecium and Pyrenula |
Ertz & Diederich
2015 |
|
|
|
Melaspilea s. lat. |
amarkantakensis S. Joseph & G. P. Sinha: on Pertusaria
amarkantakana cupularis Müll. Arg.: on Pyrenula epigena Müll. Arg.: on Reimnitzia santensis epigraphella (Nyl.) Müll. Arg.: on Acanthothecis consocians galligena Zhurb. & I. Zhdanov.: on Pertusaria insitiva Stirt.: on Pertusaria leioplaca leciographoides Vouaux: on Verrucaria lekae Brackel & Kalb: on Sarcographa tucumana Flakus, Etayo & Kukwa: on Pertusaria |
Clauzade et al. 1989 Ertz & Diederich
2015 Flakus et al. 2014 Joseph & Sinha
2015 Kalb et al. 2012 Zhurbenko &
Zhdanov 2013 |
|
|
|
Melaspileopsis |
cf.
diplasiospora (Nyl.) Ertz & Diederich: on Graphis
elegans |
Ertz & Diederich
2015 |
|
cf. diplasiospora (Ertz 16625): nrLSU: KP456166; mtSSU: KP456190 cf. diplasiospora (Ertz 16624): nrLSU: KP456165; mtSSU: KP456189 cf. diplasiospora (Ertz 16247): nrLSU: KP456164; mtSSU: KP456188 |
|
Taeniolella s. lat. |
atricerebrina Hafellner: on Tephromela beschiana Diederich: on Cladonia [= Ameroconium
cladoniae U. Braun & Zhurb.] breviuscula (Berk. & M. A. Curtis) S. Hughes: facultative
lichenicolous caespitosa M. S. Cole & D. Hawksw.: on Pertusaria chrysothricis Diederich: on Chrysothrix candelaris cladinicola Alstrup: on Cladonia subgen. Cladina delicata M. S. Christ. & D. Hawksw.: on various lichens diederichiana Etayo & Calat.: on Placopsis friesii (Hepp) Hafellner: on Strigula stigmatella hawksworthiana Heuchert, Ertz & Common: on Phaeographis ionaspisicola Alstrup & E. S. Hansen: on Ionaspis odora phaeophysciae D. Hawksw.: on Phaeophyscia
orbicularis, possibly also on Physcia
and Physconia pseudocyphellariae Etayo: on Pseudocyphellaria punctata M. S. Christ. & D. Hawksw.: on Graphis,
more rarely on Arthonia, Fissurina, Pertusaria and Phaeographis pyrenulae
Heuchert & Diederich: on Pyrenula rolfii Diederich & Zhurb.: on Cetraria and Cetrariella santessonii Etayo & Heuchert: on Roccellina cerebriformis strictae Alstrup: on Cladonia stricta toruloides Heuchert & Diederich: on Thelotrema trapeliopseos Diederich: om Trapeliopsis verrucosa M. S. Christ. & D. Hawksw.: on Arthonia
pruinata |
Alstrup 1993b Alstrup & Hansen
2001 Alstrup &
Hawksworth 1990 Cole &
Hawksworth 2001 Diederich 1990 Diederich 1992b Diederich &
Zhurbenko 1997 Diederich &
Zhurbenko 2001 Ertz et al. 2016 Hafellner 2007 Hawksworth 1979a Zhurbenko &
Alstrup 2004 |
|
hawksworthiana (Common 9199B): nrLSU: KX244970;
mtSSU: KX244984 punctata (Diederich 16040):
nrLSU: KX244973 punctata (Cezanne-Eichler 7279): nrLSU: KX244974 punctata (Ertz 17390): nrLSU: KX244972; mtSSU: KX244986 punctata (Diederich 16714): nrLSU: KX244971; mtSSU: KX244985 pyrenulae (Diederich 17075): mtSSU: KX244988 tortuloides (Diederich 17048): mtSSU: KX244990 tortuloides (Diederich 17047): mtSSU: KX244989 |
|
Stictographa |
dirinariicola
Diederich & Ertz: on Dirinaria picta lentiginosa (Leight.) Müll. Arg.: on Phaeographis dendritica |
Ertz & Diederich
2015 Diederich et al.
2017 |
|
lentiginosa (van den Boom 47621): nrLSU: KP456171; mtSSU: KP456195 |
|
Capnodiales |
|
|
|
|
|
Cladosporiaceae |
|
|
|
|
|
Cladosporium |
antarcticum K. Schub., Crous & U. Braun: isolated from Caloplaca
regalis, probably lichenicololous arthoniae M. S. Christ. & D. Hawksw.: on corticolous lichens
with Trentepohlia [not Cladosporium; Taeniolella-like] licheniphilum Heuchert & U. Braun:
on Cetraria, Glypholecia, Hypogymnia, Lecanora, Peltigera,
Pertusaria, Phaeophyscia, Physcia, Physconia Thamnolia
and Xanthoria |
Bensch et al. 2012 Diederich 1989 Hawksworth 1979a Heuchert & Braun
2006 Zhurbenko 2012 |
antarcticum: CBS 690.92 licheniphilum: CPC 13224; CBS 125990 |
antarcticum (CBS 690.91): ITS:
EF679334; TEF1: EF679405; histone H3: EF679636; cmdA: EF679560; act: EF679484 licheniphilum (CPC
13224): ITS: NR_119847 licheniphilum (CBS
125990): ITS: HM148111; TEF1: HM148355; act: HM148600 |
|
Verrucocladosporium |
dirinae K. Schub., Aptroot & Crous: on Dirina
massiliensis roccellae U. Braun, Diederich & Heuchert: on Roccella |
Braun et al. 2009 Crous et al. 2007 |
dirinae: CBS 112794 |
dirinae (CBS
112794): ITS: EU040244 |
|
Dissoconiaceae |
|
|
|
|
|
Ramichloridium |
cladoniicola U. Braun & Heuchert: on Cladonia portentosa tropicum
U. Braun &
Diederich: on crustose lichen with Trentepohlia |
Braun et al. 2009 Diederich et al.
2017 |
|
|
|
Mycosphaerellaceae |
|
|
|
|
|
Pseudocercospora |
lichenum (Keissl.) D. Hawksw.: on Haematomma cismonicum |
Etayo 2002 Hawksworth 1979a |
|
|
|
Sphaerulina s. lat. |
dolichotera (Nyl.) Vouaux: on Collemataceae [probably a synonym of
Merismatium decolorans] dubiella (Nyl.) W. Wats.: on crustose lichen intermedia Vouaux: on Leptogium [probably a synonym of Merismatium
decolorans] lepidiotae (Anzi) Vain.: on Fuscopannaria praetermissa |
Clauzade et al. 1989 |
|
|
|
Wernerella |
acarosporae
Etayo: on Acarospora nodulosa |
Etayo 2010c Navarro-Rosinés et
al. 1998d |
|
|
|
Teratosphaeriaeae |
|
|
|
|
|
Austrostigmidium |
mastodiae
Pérez-Ortega & Garrido-Benavent: on Mastodia
tessellata |
Pérez-Ortega et al.
2015 |
|
mastodiae (MA: 18210): nrLSU: KP282858 mastodiae (MA: 18215): nrLSU: KP282859 mastodiae (MA: 18214): nrLSU: KP282860 |
|
Xanthoriicola |
physciae (Kalchbr.) D. Hawksw.: on Xanthoria |
Hawksworth 1979a Hawksworth &
Punithalingam 1973 Ruibal et al. 2011 |
|
physciae (CR4): nrLSU: JN040487 physciae (CR10): nrLSU: JN040488 physciae (CR20): nrLSU: JN040489 physciae (CR21): nrLSU: JN040490 physciae (CR22): nrLSU: JN040491 physciae (CR23): nrLSU: JN040492 physciae (CR44): nrLSU: JN040493 physciae (CR45): nrLSU: JN040494 physciae (CR50): nrLSU: JN040495 physciae (CR51): nrLSU: JN040496 physciae (CR58): nrLSU: JN040497 physciae (Diederich
16713): nrLSU: KF176965; mtSSU: KF176990 |
|
Collemopsidiales |
|
|
|
|
|
Xanthopyreniaceae |
|
|
|
|
|
Collemopsidium |
cephalodiorum
(Triebel & Grube) Grube: on Pilophorus cephalodiorum |
Grube 2005 |
|
|
|
Didymellopsis |
collematum (J. Steiner) Grube & Hafellner: on Collema latitans (Nyl.) Clem. & Shear: on Gonohymenia and Lempholemma nephromatis Zhurb. & Etayo: on Nephroma cellulosum perigena (Nyl.) Grube & Hafellner: on Placidium
squamulosum pulposi (Zopf) Grube & Hafellner: on Collema and Leptogium viridireagens Etayo: on Leptogium |
Etayo 2017 Grube &
Hafellner 1990 Zhurbenko et al.
2015b |
|
|
|
Zwackhiomyces |
argentinae (Räsänen) D. Hawksw. & V. Atienza: on
Acarospora aspiciliae Halıcı & Candan: on Aspicilia berengerianus ( calcariae (Flagey) Hafellner & Nik. Hoffm.: on Aspicilia calcisedus Cl. Roux: on Verrucariaceae cervinae Calat., Triebel & Pérez-Ortega: on Acarospora cladoniae (C. W. Dodge) Diederich: on Cladonia coepulonus ( diederichii D. Hawksw. & Iturr.: on Cladonia dispersus (Körb.) Triebel & Grube: on Protoblastenia
rupestris echinulatus Brackel: on Physconia euplocinus Hafellner, Grube & Egan: on Speerschneidera
euploca heppiae van den Boom: on Heppia arenacea kantvilasii S. Y. Kondr.: on Parmotrema kiszkianus D. Hawksw. & Miadl.: on Peltigera lacustris (Arnold) Orange: on Ionaspis lacustris lecanorae (Stein) Nik. Hoffm. & Hafellner: on Lecanora lithoiceae (B. de Lesd.) Hafellner & V. John: on
Verrucaria macrosporus Alstrup & Olech: on Protopannaria martinatianus ( melanohaleae
Etayo & van den Boom: on Melanohalea
exasperata namibiensis Diederich & M. Schultz: on Psorotichia peltigerae Miadl. & Alstrup: on Peltigera physciicola Alstrup: on Physcia caesia polischukii Darmostuk & Khodos.: on Bacidia rolfii (Etayo) Etayo: on Candelina sipmanii Diederich & Zhurb.: on Phaeorrhiza sareptana socialis (Körb.) Cl. Roux: on Clauzadea, perhaps also on
Bacidia bagliettoana solenopsorae van den Boom: on Solenopsora holophaea sphinctrinoides (Zwackh) Grube & Hafellner: on Lecanora
campestris turcicus Kocakaya, Halıcı & Aksoy: on Physcia
magnussonii |
Alstrup 1993a Alstrup & Olech
1993 Aptroot et al. 1997 Diederich &
Zhurbenko 2009 Etayo 2010b Grube &
Hafellner 1990 Halıcı
& Candan 2009 Harris 1995 Hawksworth & Atienza
1994 Hawksworth &
Iturriaga 2006 Hoffmann &
Hafellner 2000 Kocakaya et al. 2011 Khodosovtsev &
Darmostuk 2017 Kondratyuk 1996b Miadlikowska &
Alstrup 1995 Orange 2002 Roux 2014 van den Boom &
Etayo 2014 |
|
|
|
Dothideales |
|
|
|
|
|
Aureobasidiaceae |
|
|
|
|
|
Pseudoseptoria |
usneae (Vouaux) D. Hawksw.: on Usnea |
Hawksworth 1981 |
|
|
|
Dothideaceae |
|
|
|
|
|
Plowrightia |
mereschkowskyi Vouaux: on Aspicilia hispida |
Clauzade et al. 1989 |
|
|
|
Lichenotheliales |
|
|
|
|
|
Lichenotheliaceae |
|
|
|
|
|
Lichenothelia |
convexa Henssen: facultative lichenicolous over siliceous
rocks renobalesiana D. Hawksw. & V. Atienza: on endolithic
Verrucariaceae (Bagliettoa, Thelidium, Verrucaria) over calcareous
rocks spiralispora Etayo [as ‘spiratispora’]:
on Acarospora sphaerospora |
Atienza &
Hawksworth 2008 Ertz et al. 2014b Kocourková &
Knudsen 2011b |
rugosa: Cultures obtained by Ertz et al. (2014) |
convexa (Diederich 17491): nrLSU: KF176962; mtSSU: KF176988 convexa (L1609): nrSSU: KC015086 convexa (L1608): nrSSU: KC015085 convexa (L1607): nrSSU: KC015084 convexa (L1606): nrSSU: KC015083 convexa (L1702): nrLSU: KC015072 rugosa (Ertz 16065): nrLSU: KF176964 rugosa (Diederich 17310): nrLSU:
KF176963; mtSSU:
KF176989 sp.
(L986): nrSSU: KC015089 sp.
(L985): nrSSU: KC015088 sp.
(L984): nrSSU: KC015087 |
|
Lichenostigma s. lat. (see under
Lichenostigmatales for Lichenostigma s.str.) |
amplum Calat. & Hafellner: on Buellia anatolicum Halıcı & Kocakaya: on brown Acarospora bolacinae Nav.-Ros., Calat. & Hafellner: on Caloplaca
bolacina canariense Etayo & van den Boom: on Aspicilia calcarea cosmopolites Hafellner & Calat.: on Xanthoparmelia dimelaenae Calat. & Hafellner: on Dimelaena oreina diploiciae Calat., Nav.-Ros. & Hafellner: on Diploicia
subcanescens elongatum Nav.-Ros. & Hafellner: on Aspicilia and Lobothallia epipolinum Nav.-Ros., Calat. & Hafellner: on Diplotomma
epipolium epiporpidiae S. Y. Kondr., L. Lőkös & J.-S. Hur: on Porpidia epirupestre Pérez-Ortega & Calat.: on Pertusaria pertusa episulphurella Etayo & van den Boom: on Lecanora sulphurella gracile Calat., Nav.-Ros. & Hafellner: on Acarospora
fuscata heterodermiae S. Y. Kondr., L. Lőkös et J.-S. Hur: on Heterodermia iranicum Brackel & Valadbeigi: on Lobothallia
praeradiosa rouxii Nav.-Ros., Calat. & Hafellner: on Squamarina rupicolae Fern.-Brime & Nav.-Ros.: on Pertusaria rupicola saxicola K. Knudsen & Kocourk.: saxicolous, facultative
lichenicolous on Polysporina simplex semi-immersum Hafellner: on Buellia subradians Hafellner, Calat. & Nav.-Ros.: on Acarospora supertegentis Ihlen & R. Sant.: on Aspicilia supertegens svandae Vondrák & Šoun: on Acarospora cervina triseptatum Halıcı & D. Hawksw.: on Aspicilia |
Calatayud &
Barreno 2003 Calatayud & Navarro-Rosinés
2000 Calatayud et al.
2002 Fernández-Brime et
al. 2010 Hafellner &
Calatayud 1999 Halıcı
& Hawksworth 2007 Halıcı et
al. 2009b Ihlen 2004 Kondratyuk et al.
2013 Kondratyuk et al.
2017 Knudsen &
Kocourková 2010b van den Boom &
Etayo 2006 Vondrák & Šoun
2007 |
cf. elongatum: Cultures obtained
by Ertz et al. (2014) |
Lichenotheliaceae sp.
(Ertz 17540): nrLSU:
KF176960; mtSSU: KF176986 Lichenotheliaceae sp.
(Ertz 17457): nrLSU: KF176959; mtSSU:
KF176985 Lichenotheliaceae sp.
(Ertz 16455): nrLSU: KF176958;
mtSSU: KF176984 Lichenotheliaceae |